Penalties, which have troubled Harvard all season, became a problem again on Saturday. The Crimson tallied 11 penalties for 79 yards, including the two which put Flesher’s would-be game-winning field goal out of reach.

Harvard’s red zone became a familiar place for the Princeton offense in the first quarter. Within the first 15 minutes, the Tigers found themselves inside the 20-yard line on three separate occasions—every one of their first-quarter drives.

On the first drive of the game, Epperly drove Princeton down 75 yards for the first score. A missed two-point conversion gave the Tigers the early 6-0 lead just three minutes into the game.

The Crimson defense appeared to have gotten a stop on Princeton’s following drive, but a Harvard defender ran into the punter, extending the Tigers’ drive. Princeton moved all the way down to the one on the drive, but the defense came up with a goal-line stand.

When Harvard’s offense couldn’t capitalize on the turnover, the ensuing field position in Harvard territory led to another seven points for the Tigers.

But once the second quarter began, the momentum shifted. The Crimson scored 14 unanswered points to take the lead on sophomore Paul Stanton’s longest run of the year, a 60-yard sprint to the end zone that gave Harvard a 14-13 lead.

A costly mistake by Hempel led to another Princeton score. Under pressure, Hempel threw a free-for-all that defensive back Philip Bhaya plucked out of the air. Two minutes later, Princeton turned the mistake into eight points, as Epperly hit receiver Conner Kelley for a score and converted the two points afterward.

—Staff writer Samantha Lin can be reached at samantha.Lin@thecrimson.com. Follow her on Twitter @linsamnity.