Longtine said that Cotran taught students "to value the interface between science and medicine" as well as to have confidence in their personal ability to further the field of pathology.
Both colleagues emphasized Cotran's personable nature and what Fletcher called his "extraordinary appreciation of people."
Cotran sought to improve aspiring doctors while advancing their careers, Fletcher said.
"In the incredible hierarchy of Harvard Medical School, he was approachable," Longtine said.
Fletcher said Cotran's "careful attention to the needs of others" was what "inspired the most devotion in those around him."
"He wore his accomplishments with joy rather than with pride," Fletcher said.
Born in Haifa in what is now Israel, Cotran received both his undergraduate and medical degrees from the American University of Beirut. He became an HMS professor in 1960.
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